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1a
Thread-like structure in living cells carrying genetic information (10)
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7a
Queensland's capital (8)
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8a
Part of the mouth (4)
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9a
Gardener's shallow basket (4)
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10a
Rugby position (3,4)
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12a
At present — fetch Ron one (anag) (3,3,5)
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14a
Treacle (anag) — daughter of Agamemnon (7)
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16a
Bargain — short cut? (4)
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19a
Joint often used in dandling (4)
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20a
One of Shakespeare's tragedies (4,4)
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21a
Short leather trousers with braces (10)
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1d
Measure of gold purity (5)
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2d
Saviour (7)
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3d
An average (4)
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4d
Young plant (8)
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5d
One of the gifts brought by the Magi (5)
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6d
Border, Scotch or Welsh sheepdog (6)
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11d
A Native American people — coke here (anag) (8)
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12d
Crime, such as murder or arson (6)
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13d
Flummox (7)
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15d
Greek island, base of the Minoan civilisation (5)
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17d
Without bells and whistles (5)
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18d
Rainfall measurement unit (4)